Based on Plate 5 – “Le Grand Toucan a gorge orange” of “Histoire naturelle des oiseaux de paradis et des rolliers, suivie de celles des toucans et des barbus. Paris, Denné le jeune & Perlet, (1801–1806). Volume 2″ by Jacques Barraband (1767-1809).
The colorful and large bill, which in some large species measures more than half the length of the body, is the hallmark of toucans. Despite its size, the toucan’s bill is very light, being composed of bone struts filled with spongy tissue of keratin between them. The bill has forward-facing serrations resembling teeth, which historically led naturalists to believe that toucans captured fish and were primarily carnivorous; today it is known that they eat mostly fruit.